The primitive crocodiles, which lived 100m years ago, were good swimmers but were also capable of galloping
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Three of the crocodiles are new species and include Kaprosuchus saharicus, a 6.5m-long beast with three sets of dagger-like tusks and an armoured snout for ramming its prey.
Another species, Laganosuchus thaumastos, was of similar length but had a pancake-flat head and is thought to have lurked in rivers with its jaws open, waiting for unsuspecting fish to pass.
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The most striking feature the beasts have in common was revealed by their bone structure, which suggests they were efficient swimmers but that when they clambered ashore they were also capable of galloping across the plains. Modern crocodiles crawl on their bellies because their legs sprawl out to the side.
"My African crocs appeared to have had both upright, agile legs for bounding overland and a versatile tail for paddling in water," writes Paul Sereno, a palaeontologist at the University of Chicago, in National Geographic Magazine. "These species open a window on a croc world completely foreign to what was living on northern continents."
The third new species, Araripesuchus rattoides, was only a metre long and probably used a pair of buckteeth in its lower jaw to dig for grubs.
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Most of the fossils were found near the site where, in 2001, Sereno uncovered a 12m-long crocodile that lived 110m years ago. The beast, nicknamed SuperCroc, weighed around eight tonnes. The latest fossils are described in the journal ZooKeys.
"We were surprised to find so many species from the same time in the same place," said Hans
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The expedition was sponsored by National Geographic, which airs a documentary about the discoveries, When Crocs Ate Dinosaurs, at 5pm on Sunday 20 December on the Nat Geo Wild channel.
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